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About Traditional Art / Professional Brian ChooMale/Australia Group :iconprehistory-alive: Prehistory-Alive
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Moythomasia lineata by Gogosardina
Moythomasia lineata
2013+15, acrylics, digital and photography

Image for = Choo, B. (2015) A new species of the Devonian actinopterygian Moythomasia from Bergisch Gladbach, Germany, and fresh observations on M. durgaringa from the Gogo Formation of Western Australia, Journal of Vertebrate Paleontology. DOI:10.1080/02724634.2015.952817

link =…

ca. 380 million years ago, Late Devonian (early Frasnian), Oberer Plattenkalk, Bergisch-Gladbach, Nordrheine-Westfalen, Germany.

In a shallow sea over what will become Germany, a 20 cm long actinopterygian (Moythomasia lineata) swims in front of a shoal of smaller cousins (Moythomasia nitida) along with a tetrapodomorph (Latvius niger) and a trio of pelagic crustaceans (Montecaris strunensis)

Moythomasia lineata = an ancient actinopterygian fish named and described in the above paper by yours truly. Based on material originally collected in the 1960s and currently house in Naturhistoriska riksmuseet, Stockholm. Known from three incomplete large adults and and least 8 small juveniles. Larger specimens (including the eventual holotype) were initially described as M. cf.striata by Jessen (1968), a scale-taxon from Wildungen while the juvies were misidentified as the contemporaneous M. nitida. When I visited the collections in 2007, I determined that what we had was a new species of Moythomasia, showing that you can make discoveries without getting your hands dirty by simply snooping around museum collections. M. lineata differs from M. nitida (plus the Aussie M. durgaringa) in have a completely linear bone and scale ornament, without the little pits you find in the other species. It's also more elongate that other completely known members of the genus. The scales of the genuine M. striata have a similar ornament to lineata but a different shape and it is not clear that they belong to Moythomasia in the first place.   

Moythomasia nitida = the most common fossil fish at Bergisch-Gladbach. Originally named by Gross in 1953, however when numerous complete specimens were described by Jessen i(1968), it was the first time such an ancient ray-finned fish had been described in such detail. 

Latvius niger = A small osteolepid tetrapodomorph.

Montecaris strunensis = a phyllocarid, malacostracan crustaceans with a bivalved carapace. Living phyllocarids (leptostracans) are less than an inch long, but Devonian archaeostracan examples could grow much bigger. Montecaris was a highly successful genus of the Middle-Late Devonian with a wide distribution (Europe, Canada, Australia), typically ranging in size from 5 - 25 cm in length (with rare examples approaching 2 feet).
Yi qi. Dragon of the Daohugou by Gogosardina
Yi qi. Dragon of the Daohugou
2015, digital, pencils and photography 2015.

(edit 10/05/2015 = diminished the excessive wing-creasing on the foreground animal)

ca.164 million years ago, Middle-Late Jurassic boundary (Callovian-Oxfordian), Tiaojishan Formation (Daohugou Biota), Hebei, China

As dusk approaches, the bantam-sized scansoriopterygid dinosaur Yi qi begins to fastidiously preen the feathers in preparation for a twilight foraging session.

Yi qi ("strange wing" in mandarin) is one incredible animal.

The exquisite fossil holotype of Yi hails from Hebei Province, in northern China. Extensive feather impressions are preserved around most of the body. But the most remarkable things are the hands. The three digits are greatly elongated (as in other scansoriopterygids like Epidendrosaurus). But there is also an enormous prong-like structure (styliform element) sticking out of each wrist. Preserved adjacent to these are broad patches of naked skin, devoid of scales or feathers.

The most likely explanation =  The three elongated fingers and the extra bony strut supported an aerodynamic flight membrane. Yi was a theropod with draconic-wings. The first ever dinosaur in the fossil record with a membranous rather than a feathery flight apparatus. It is unclear if Yi was capable of powered flight or simply glided (the authors of the paper suspect the latter).

Yi is the latest find from the Daohugou biota, a series of mid-Jurassic sites in Hebei, Liaoning and Inner Mongolia with identical super-preservation as the more famous Jehol biota of Cretaceous Liaoning. The discovery of this fauna destroyed the so-called temporal paradox (an objection to the dino-to-bird hypothesis), based on the supposed absence of bird-like dinosaurs before the traditional "first bird" Archaeopteryx. The Daohugou represents a community of feathered dinosaurs over 10 million years older than Archie.  

 Yi shared it's world with a host gliding and flying vertebrates = avialian dinosaurs (Anchiornis, Xiaotingia), pterosaurs (Darwinopterus, Jeholopterus) and gliding mammals (Volaticotherium). Vertebrate evolution seemed to have been cashing in on the flight-craze during the Jurassic of China, with numerous clades independently putting representatives in the air.

Ultimately Yi demonstrates that the transition from dinosaur-to-bird was not straightforward. Aerial adaptations evolved multiple times among the maniraptorans with a great deal of early innovation in flight. Yi's strange wing did not lead to birds, it was happily doing it's own thing.

Note that the colouration is based on taking actual fossil melanosome data at face value. According to personal communications with Xu Xing, Yi probably had very dark feathers over the head and a more neutral colouration over the rest of the body.


  • Xing Xu, Xiaoting Zheng, Corwin Sullivan, Xiaoli Wang, Lida Xing, Yan Wang, Xiaomei Zhang, Jingmai K. O’Connor, Fucheng Zhang & Yanhong Pan. 2015. A bizarre Jurassic maniraptoran theropod with preserved evidence of membranous wings. Nature doi:10.1038/nature14423

We made this holoscape with pics we downloaded off Google images - spared no expense.…


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Brian Choo
Artist | Professional | Traditional Art
An Australian vertebrate palaeontologist. I mainly work on Silurian-Devonian fishes, especially basal actinopterygians.

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HUBLERDON Featured By Owner 3 hours ago  Hobbyist General Artist
Hello, Brian!

I was a big fan of Spec World in my youth, and I've recently heard you created some sort of bizarre "Caripoo" in-joke in the community. But there's nothing on the net explaining what it's supposed to be! Can you explain, please? :)

g4321g5 Featured By Owner 6 days ago  Hobbyist General Artist
Happy Belated birthday! :D
Helixdude Featured By Owner Nov 21, 2015  Hobbyist Traditional Artist
Happy Birthday!!
Saberrex Featured By Owner Nov 21, 2015  Hobbyist General Artist
Happy birthday. 
Dontknowwhattodraw94 Featured By Owner Nov 21, 2015  Hobbyist Traditional Artist
Happy birthday!
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